Okay,
1. Put values on the tower attributes, meters are hard to judge. A number within the meter is fine.
2. Speed increase/decrease functions.
3. Possibly alternate paths that different enemies use (air is overused - use water paths maybe? Would go well with the atmosphere you have going).
4. One tower with a relocate ability would be nice.
I might think of more later, but I love the art style on this game and want it to go far. Keep it up!
Awesome game. Could use a few more effects within the cave - the outside looks amazing, the inside is pretty dull in some areas. Loved the storyline too. 4/5
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