wildbill, no need to upgrade only damage, for any combo to work damage must be max at every part of that combo, for example: Nuke- both cannons must have max damage and also rocket launcher, you can upgrade their range to max or their rate ;)
I had that taactic aat beggining, but then from 12th level on I went by destroying their weakest cell and getting 1 for myself, then destroying any they populated. It's a bit bugging as you constantly lose your cells, but it does the trick
Reproduction and speed, well also attack, so basicly a destructing tank that moves extra fast.
Had problems at start due to lag...5/5, one of my favourite games =)