[Cursed Treasure: Don't Touch My Gems!] A question about skill benefits.

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When ever i spend a skill point in a tree, lets say Orcs in this scenario, i get 1% fire rate bonus. When ever i upgrade Undead, i get 1% range bonus. Do these bonuses only apply to their own “race”, or do they apply to all the towers?

Tl;dr
Can the fire rate bonus from putting skills in the Orc tree apply to Crypts and Temples?

 
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It applies to all of your towers.

 
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Are you sure? I think its only for the specific tower upgraded section.

 
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It’s definitely to all towers. 20% damage bonus from Temple upgrading becomes quite noticeable on your Crypts and Dens.