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myninth it can go through two separate bloons. so if there were 2 reds next to each other, it could pop them both. nothing except the upgraded mortar can pop through two bloon LAYERS at once
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This is in response to Myninthpassword's queries: Yes, the Dart Monkey's second upgrade allows him to pop two [outer layer] bloons. Basically, what you are asking refers to how they describe "layers" of bloons. The only tower which can "pop" multiple layers of a bloon at a time is the Mortar Tower. (Though there is the Super Monkey Storm, which is an ability available with with your first fully upgraded Monkey Beacon).
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This game kind of gives me a way of thinking that a TD game is like a digestive system to use the right organs on each food (bloons) is essential for winning anyway but there's also a lot of difference anyway, They're still the same at that point.
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I'm totally not cheating by buying high-tier upgrades early. (I upgrade a tower, and before the upgrade animation finishes, I click on another tower and upgrade a higher tier.)